@define-mixin dark-theme-mutations {
  :root {
    /* Repeat variable definitions as CSS variables */
    --background-color: $background-color;
    --page-background-color: $page-background-color;
    --foreground-color: $foreground-color;
    --base-color: $base-color;
    --primary-light-color: $primary-light-color;
    --success-light-color: $success-light-color;
    --danger-light-color: $danger-light-color;
    --warning-light-color: $warning-light-color;
    --destroyed-content-color: $destroyed-content-color;
    --header-color: $header-color;
    --header-field-color: $header-field-color;
    --header-secondary-color: $header-secondary-color;
    --header-admin-color: $header-admin-color;
    --block-header-color: $block-header-color;
    --block-header-link-text-color: $block-header-link-text-color;
    --block-header-light-color: $block-header-light-color;
    --block-header-light-link-text-color: $block-header-light-link-text-color;
    --border-color: $border-color;
    --media-box-color: $media-box-color;
    --link-color: $link-color;
    --link-hover-color: $link-hover-color;
    --fave-color: $fave-color;
    --vote-up-color: $vote-up-color;
    --vote-down-color: $vote-down-color;
    --hide-color: $hide-color;
    --assistant-color: $assistant-color;
    --unread-message-color: $unread-message-color;
    --tag-normal-color: $tag-normal-color;
    --tag-category-rating-color: $tag-category-rating-color;
    --tag-category-spoiler-color: $tag-category-spoiler-color;
    --tag-category-origin-color: $tag-category-origin-color;
    --tag-category-oc-color: $tag-category-oc-color;
    --tag-category-error-color: $tag-category-error-color;
    --tag-category-character-color: $tag-category-character-color;
    --tag-category-content-official-color: $tag-category-content-official-color;
    --tag-category-content-fanmade-color: $tag-category-content-fanmade-color;
    --tag-category-species-color: $tag-category-species-color;
    --tag-category-body-type-color: $tag-category-body-type-color;
    --commission-category-color: $commission-category-color;
    --site-notice-color: $site-notice-color;
    --image-overlay-color: $image-overlay-color;
    --image-overlay-background-color: $image-overlay-background-color;
    --text-light-color: $foreground-color;

    --meta-color: hsl(from $meta-color h calc(s - 8) calc(l + 6));
    --meta-border-color: hsl(from $meta-color h calc(s - 15) calc(l + 25));

    --comment-color: var(--link-hover-color);
    --link-light-color: var(--foreground-color);

    --primary-color: hsl(from $primary-light-color h calc(s + 4) calc(l + 5));
    --success-color: hsl(from $success-light-color h s calc(l + 4));
    --danger-color: hsl(from $danger-light-color h s calc(l + 5));
    --warning-color: hsl(from $warning-light-color h s calc(l + 5));

    --header-hover-color: hsl(from $header-color h s calc(l - 8));
    --header-field-hover-color: hsl(from $header-color h s calc(l - 12));
    --header-secondary-hover-color: hsl(from $header-secondary-color h s calc(l - 4));

    --header-admin-hover-color: hsl(from $header-admin-color h s calc(l - 5));

    --block-header-hover-color: hsl(from $block-header-color calc(h - 3) calc(s + 4) calc(l - 4));
    --block-header-link-text-hover-color: hsl(from $block-header-link-text-color calc(h + 6) calc(s - 20) calc(l - 3));

    --block-header-light-hover-color: hsl(from $block-header-light-color calc(h - 4) calc(s + 10) calc(l - 4));
    --block-header-light-link-text-hover-color: hsl(from $block-header-light-link-text-color calc(h + 8) calc(s - 10) calc(l - 2));

    --media-box-hover-color: hsl(from $media-box-color h s calc(l - 4));
    --media-box-header-link-text-hover-color: hsl(from $link-color h calc(s - 18) calc(l - 3));
    --media-box-header-link-text-color: $link-color;
    --media-box-header-link-text-hover-color: hsl(from $link-color h s calc(l - 10));

    --link-dark-color: hsl(from $link-color h calc(s - 15) calc(l + 40));
    --link-dark-hover-color: hsl(from $link-hover-color h s calc(l + 40));

    --image-show-link-color: hsl(from $link-color h s calc(l + 40));
    --image-show-link-hover-color: hsl(from $link-hover-color h s calc(l + 40));

    --image-overlay-color: $foreground-color;
    --image-overlay-background: hsl(from $background-color h s calc(l - 30));

    --assistant-border-color: hsl(from $assistant-color h s calc(l + 8));

    --admin-links-color: hsl(from $vote-down-color h calc(s - 50) calc(l - 28));
    --admin-links-hover-color: hsl(from $vote-down-color h calc(s - 20) calc(l - 20));

    --background-odd-color: hsl(from $background-color h s calc(l - 3));
    --background-even-color: $background-color;

    --success-odd-color: hsl(from $success-light-color h calc(s - 2) calc(l - 5));
    --success-even-color: hsl(from $success-light-color h calc(s + 2) calc(l - 2));
    --warning-odd-color: hsl(from $warning-light-color h calc(s + 10) calc(l - 15));
    --warning-even-color: hsl(from $warning-light-color h calc(s + 7) calc(l - 7));
    --danger-odd-color: hsl(from $danger-light-color h calc(s - 2) calc(l - 11));
    --danger-even-color: hsl(from $danger-light-color h s calc(l - 6));

    --foreground-half-color: hsl(from $background-color h s calc(l + 50));

    --sparkline-color: hsl(from $base-color h s calc(l + 5));

    --input-text-color: $foreground-color;
    --input-color: var(--meta-color);
    --input-border: var(--meta-border-color);
    --input-color-active: hsl(from $meta-color h s calc(l + 5));
    --input-border-active: hsl(from $base-color h calc(s - 10) calc(l + 2));

    --tag-normal-background: hsl(from $tag-normal-color h s calc(l - 29));
    --tag-normal-border: hsl(from $tag-normal-color h s calc(l - 18));
    --tag-category-rating-background: hsl(from $tag-category-rating-color h s calc(l - 35));
    --tag-category-rating-border: hsl(from $tag-category-rating-color h calc(s - 10) calc(l - 20));
    --tag-category-spoiler-background: hsl(from $tag-category-spoiler-color h s calc(l - 34));
    --tag-category-spoiler-border: hsl(from $tag-category-spoiler-color h s calc(l - 23));
    --tag-category-origin-background: hsl(from $tag-category-origin-color h s calc(l - 40));
    --tag-category-origin-border: hsl(from $tag-category-origin-color h calc(s - 28) calc(l - 22));
    --tag-category-oc-background: hsl(from $tag-category-oc-color h s calc(l - 33));
    --tag-category-oc-border: hsl(from $tag-category-oc-color h s calc(l - 15));
    --tag-category-error-background: hsl(from $tag-category-error-color h calc(s - 6) calc(l - 38));
    --tag-category-error-border: hsl(from $tag-category-error-color h s calc(l - 22));
    --tag-category-character-background: hsl(from $tag-category-character-color h s calc(l - 33));
    --tag-category-character-border: hsl(from $tag-category-character-color h s calc(l - 20));
    --tag-category-content-official-background: hsl(from $tag-category-content-official-color h calc(s - 2) calc(l - 29));
    --tag-category-content-official-border: hsl(from $tag-category-content-official-color h s calc(l - 20));
    --tag-category-content-fanmade-background: hsl(from $tag-category-content-fanmade-color h s calc(l - 40));
    --tag-category-content-fanmade-border: hsl(from $tag-category-content-fanmade-color h calc(s - 10) calc(l - 20));
    --tag-category-species-background: hsl(from $tag-category-species-color h s calc(l - 35));
    --tag-category-species-border: hsl(from $tag-category-species-color h calc(s - 10) calc(l - 20));
    --tag-category-body-type-background: hsl(from $tag-category-body-type-color h s calc(l - 50));
    --tag-category-body-type-border: hsl(from $tag-category-body-type-color h calc(s - 10) calc(l - 37));

    --commission-category-background: hsl(from $commission-category-color h s calc(l - 36));
    --commission-category-border: hsl(from $commission-category-color h s calc(l - 25));

    --site-notice-link-color: hsl(from $block-header-link-text-color h calc(s - 20) calc(l + 8));
    --site-notice-link-hover-color: hsl(from $block-header-link-text-color h calc(s - 20) l);

    --button-text-color: $foreground-color;
    --button-background-color: rgba($meta-color, 0.2);
    --button-hover-background-color: var(--input-color-active);
    --button-hover-border-color: var(--input-border-active);
    --button-border-color: var(--meta-border-color);

    --button-primary-background-color: $primary-light-color;
    --button-success-background-color: $success-light-color;
    --button-warning-background-color: $warning-light-color;
    --button-danger-background-color: $danger-light-color;

    --button-primary-border-color: hsl(from $primary-light-color h calc(s + 22) calc(l + 14));
    --button-success-border-color: hsl(from $success-light-color h calc(s + 10) calc(l + 9));
    --button-warning-border-color: hsl(from $warning-light-color h calc(s + 10) calc(l + 12));
    --button-danger-border-color: hsl(from $danger-light-color h calc(s + 6) calc(l + 18));

    --button-primary-hover-background: hsl(from $primary-light-color h calc(s + 5) l);
    --button-primary-hover-border: hsl(from $primary-light-color h calc(s + 5) l);
    --button-success-hover-background: hsl(from $success-light-color h s calc(l - 5));
    --button-success-hover-border: hsl(from $success-light-color h calc(s + 10) calc(l + 4));
    --button-warning-hover-background: hsl(from $warning-light-color h s calc(l - 6));
    --button-warning-hover-border: hsl(from $warning-light-color h calc(s + 10) calc(l + 6));
    --button-danger-hover-background: hsl(from $danger-light-color h s calc(l - 5));
    --button-danger-hover-border: hsl(from $danger-light-color h calc(s + 6) calc(l + 13));

    --downvote-disabled-color: hsl(from $vote-down-color h calc(s - 10) calc(l - 10));
    --dnp-warning-hover-color: hsl(from $vote-down-color h s calc(l + 10));
    --poll-form-label-background: hsl(from $border-color h s calc(l + 8));
    --tag-dropdown-hover-background: hsl(from $meta-color h s calc(l - 4));
  }
}